Name: Arianny Pimentel Storari
Type: MSc dissertation
Publication date: 28/02/2020
Advisor:

Namesort descending Role
Frederico Falcão Salles Co-advisor *
Taissa Rodrigues Marques da Silva Advisor *

Examining board:

Namesort descending Role
Frederico Falcão Salles Co advisor *
Hermínio Ismael de Araújo Júnior External Examiner *
Marcelo Teixeira Tavares Internal Examiner *
Mírian Liza Alves Forancelli Pacheco External Alternate *
Taissa Rodrigues Marques da Silva Advisor *

Summary: Among the groups of insects, mayflies stand out in the
Crato Formation, Araripe Basin. A newly discovered adult
individual from the family Oligoneuriidae is here described:
Incogemina nubila gen. et sp. nov. and new subfamily. Also, a
previously described “oligoneuriid” specimen from the same
unit is here reviewed, redescribed and considered as belonging to
the family Hexagenitidae. The Incogemina bridge an important gap
between an oligoneuriid ancestor and the extant Chromarcys.
Furthermore, they demonstrate that the divergence between
Oligoneuriinae + Incogemiinae and Chromarcynae + Colocrurinae
probably occurred in South America. Additionally, we described
recently collected fossil specimens of extinct family
Hexagenitidae. Two new adults specimen were described as
Protoligoneuria limai, and the taxon diagnosis was emended,
besides the description of two more adult specimens of the
Hexagenitidae, not named. Therefore, we consider that
Cratohexagenites minor could be synonymous with P. limai,
providing new morphological information about the Crato Formation
mayfly fauna, specially the hexagenitids. Finally, we analyze a
mayfly mass mortality collected at controlled excavations, the
first of their kind for the Crato Formation. All mayflies found
belong to the Hexagenitidae, whose morphology demonstrates that
the nymphs occurred in quiet waters. The nymphs were smaller than
usual, suggesting a shallow water column, with excellent
preservation and no preferential orientation in the samples,
suggesting lack of transport. Thus, we conclude that the Crato
Formation lake experienced a seasonal high evaporation caused by
the hot climate tending to aridity.

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